Inkwater Marsh

Head left to find the first Life Cell Fragment. A spirit orb is below a log, but this area is inaccessible for now. Keep heading left, jump over the Slime (which you cannot fight yet), and jump on the platforms above the water. Use the rope to get to the other side, then climb the rock. Jump on the plant to get a life orb. Jump on the platform to the right; it will break near the rope. Jump back in and proceed to the upper area.

From the top area, head left, jump over the spikes, and pass through a billboard. Drop down and drag the rock out of the way to get a Life Cell Fragment. Collecting more fragments will upgrade your max health.

Head back to where you started and proceed to the right. As you climb the first rock, you'll see a small yellow orb. Drop down and turn left to get a small spirit light container, which can be traded for items and upgrades.

Keep heading right to find some branches blocking your way. As you return, the tree will fall with you. Head to the right corner to find another spirit light container. Jump down the platform and keep descending until you reach the bottom.

Head left to find a fireplace and get a torch. You can now fight enemies. Defeat the first enemy ahead to get an Energy Orb (acts like mana). Keep advancing to reach a locked Spirit Gate. Turn left, drop down, and break the branches to proceed. Wall jump through the rocks to find Tokk. Speak to him; he will tell you that you need two keystones to open the spirit gate and give you your first quest.

The Missing Key: Retrieve the missing keystone from the cave beyond Tokk, and he'll give you his.

After talking with Tokk, proceed leftwards and wall jump to advance. Drop down onto the wooden platform and go to the left corner. Break the branches to find a Spirit Orb.

Head back to the platform, jump over the spikes, and defeat the enemy ahead. Be careful when descending, as you can land on spikes. Keep right, drop down, and break the branches in the right corner to get a Spirit Orb. Turn left to see a closed gate. An Axolotl will engage; defeat it to proceed.

As you progress, your torch will light off. Use the blue moss to climb the walls. Jump on top of the rock and cling to the blue moss above you. Keep moving upwards to find the objective on the top-left corner of the screen. Use the blue moss to reach it and get the keystone. Head back to Tokk and talk to him to get the second keystone. Climb your way back to the spirit gate to proceed.

After passing through the gate, you will see clear water that won't hurt you. Keep advancing, and a wolf will appear. You must escape; quickly jump over the platforms and find a torch. Fight the first boss.

Boss Battle: Howl. The boss has two attack types: advancing to bite you or performing a smash attack. Hit him just two times to be safe and get some distance to dodge his attacks. If you do not attack him, he will advance through the arena.

After fighting off the Howl, return to the path and you will see an opening leading down. Jump to the left side to find a Spirit Orb.

Keep descending to reach Howl's Den. The left path is blocked by a spirit gate. The skeleton on the ground will be important later. Head right, and a little creature will run off. Keep advancing to find the first Spirit Tree. Approach it to absorb the light, and a cutscene will trigger.

After the cutscene, the little creatures will approach and talk to you. You have gained the Spirit Edge. Keep through the left path to return. As you proceed, some enemies will engage; defeat them and keep upwards. Go to the right side of the screen and break the branches to find a Spirit Shard (equipable upgrades). This one gives you the ability to attract orbs from half the screen.

Keep on the path and you will return to the location before the Spirit Tree. The little creatures, Moki, will give you a quest to find the guardian of the marsh.

Guardian of the Marsh: Find Kwolok on Kwolok's Hollow.

Drop down onto the platform where you spoke with the Moki. Defeat the enemies in the path and go to the left corner. Break the branches to find a Spirit Orb. Keep downwards, then head east to find a Spirit Well (use them to save and as warp points).

Head west and you will see a wooden platform hanging. Look right and drag a rock to the wooden platform so you can jump to the other side. Keep west to enter an area where gates will lock you in. Defeat the enemies and keep left. Drop down onto the wooden platform and head right to find a Spirit Orb.

Find a Spirit Tree on the east. Absorb its light to gain the ability to double jump. Now you can jump the wall on the left and double jump to get a Spirit Shard (grants the ability to stick and climb on walls).

Return using the path below. Go downwards to reach a place with a lever. Pull it to lower the wooden platform, then drop on it and push the rock against the laser to advance and get the keystone.

Return to where you landed in Howl's Den and jump on the wall near the gate. There you will find the next keystone. Take it and open the gate to proceed.

Double jump to cross the lake and destroy the wall on the left to find an unlocked path. Jump down onto the rock, then cling to the blue moss and jump on the walls to find a Life Cell Fragment.

Return to the previous location, head upwards, break the branches, and head east. Jump on the platforms on the water and break the wall in the corner to get a Life Cell Fragment.